Song: Males make a clear resonant "QUE-ark" that can be heard over long distances. Groups produce loud "kow kow kow" calls as greetings when reunited. Contact calls given by both sexes are clucking trills.
Geoffrey A. Keller (Macaulay Library, Cornell
# 42202); 1988 May 22; USA, Oregon; 7 km S from Charleston, South Slough Sanctuary, Coos Co; Adult; Primary Sound;