Song: Males make a clear resonant "QUE-ark" that can be heard over long distances. Groups produce loud "kow kow kow" calls as greetings when reunited. Contact calls given by both sexes are clucking trills.
Sound: Geoffrey A. Keller (
Macaulay Library, Cornell # 42202); 1988 May 22; USA, Oregon; 7 km S from Charleston, South Slough Sanctuary, Coos Co; Adult; Primary Sound;